The two lead characters appear as the robotic forms of Daft Punk and are credited as "Hero Robot No. 1" and "Hero Robot No. 2"; one wears a silver helmet and the other wears a gold one. An opening scene shows the duo driving in a 1987 Ferrari 412 with its license plate displaying "HUMAN". After passing through a Southwestern United States landscape, the duo arrives by car at a town in Inyo County, California. The town's denizens are also shown to be robots physically identical to the two main characters, but at different ages, with different clothing and alternating gender.
The pair drive to a high-tech facility where liquid latex is poured over their heads. The latex is shaped into human-like faces with the aid of prosthetic appliances and wigs. When the two leave the facility, the locals of the town are shocked by their human appearance. The duo's faces eventually melt in the sun as the townsfolk chase them. The two take cover in a public restroom where the gold robot discards his ruined mask, then encourages the reluctant silver robot to do the same. Again appearing as robots, the pair then undergo a lengthy hike across desert salt flats.
After walking for a while, the silver robot removes its jacket and reveals a switch on its back. The gold robot flips the switch, which begins a timer. When the countdown ends, the silver robot is blown to pieces. The remaining robot piles the remains of the silver robot, then continues to walk. The gold robot eventually falls to its knees and attempts to reach the switch on its own back, but to no avail. The robot removes its helmet and repeatedly slams it into the ground until the helmet shatters. Using one of the shards as a burning-glass, the robot focuses the sunlight to set its hand ablaze. The film ends as the robot, completely on fire, walks in slow motion through darkness.